See Google Scholar for up-to-date citations. Lab members in bold.

Accepted or in press

Reimer JR, Arroyo-Esquivel J, Jiang J, Scharf HR, Wolkovich EM, Zhu K, Boettiger C. Noise can create or erase long transient dynamics. Theoretical Ecology, in press.

Qiu T, …, Zhu K, …, Clark JS. Is there tree senescence? The fecundity evidence. Proceedings of the National Academy of Sciences, in press.

Qin C, Bartelme R, Chung YA, Fairbanks D, Lin Y, Liptzin D, Muscarella C, Naithani K, Peay K, Pellitier P, St. Rose A, Stanish L, Werbin Z, Zhu K. From DNA sequences to microbial ecology: Wrangling NEON soil microbe data with the neonMicrobe R package. Ecosphere, in press.

Nagy C, …, Zhu K. Harnessing the NEON data revolution to advance open environmental science with a diverse and data-capable community. Ecosphere, in press.

Jung CG, Xu X, Shi Z, Niu S, Xia J, Sherry R, Jiang L, Zhu K, Hou E, Luo Y. Warmer and wetter climate promotes net primary production in C4 grassland with additional enhancement by hay-harvesting. Ecosphere, in press.

Chowdhury S, Zhu K, Zhang Y. Mitigating greenhouse gas emissions through generative adversarial networks based wildfire prediction. To appear at the 2021 MIT “A+B” Applied Energy Symposium, Aug. 11-13, 2021, Cambridge, Massachusetts, USA.


Zhong Y, Chu C, Myers JA, Gilbert GS, Lutz JA, Stillhard J, Zhu K, Thompson J, Baltzer JL, He F, LaManna JA, Davies SJ, Aderson-Teixeira KJ, Burslem DFRP, Alonso A, Chao K-J, Wang X, Gao L, Orwig DA, Yin X, Sui X, Su Z, Abiem I, Bissiengou P, Bourg N, Butt N, Cao M, Chang-Yang C-H, Chao W-C, Chapman H, Chen Y-Y, Coomes DA, Cordell S, Oliveira AA de, Du H, Fang S, Giardina CP, Hao Z, Hector A, Hubbell SP, Janík D, Jansen PA, Jiang M, Jin G, Kenfack D, Král K, Larson AJ, Li B, Li X, Li Y, Lian J, Lin L, Liu F, Liu Y, Liu Y, Luan F, Luo Y, Ma K, Malhi Y, McMahon SM, McShea W, Memiaghe H, Mi X, Morecroft M, Novotny V, O’Brien MJ, Ouden J den, Parker GG, Qiao X, Ren H, Reynolds G, Samonil P, Sang W, Shen G, Shen Z, Song G-ZM, Sun I-F, Tang H, Tian S, Uowolo AL, Uriarte M, Wang B, Wang X, Wang Y, Weiblen GD, Wu Z, Xi N, Xiang W, Xu H, Xu K, Ye W, Yu M, Zeng F, Zhang M, Zhang Y, Zhu L, Zimmerman JK (2021). Arbuscular mycorrhizal trees influence the latitudinal beta-diversity gradient of tree communities in forests worldwide. Nature Communications, 12(1):3137.

Yang X, Zhu K, Loik ME, Sun W (2021). Differential responses of soil bacteria and fungi to altered precipitation in a meadow steppe. Geoderma, 384, 114812.

Wang F, Sanders CJ, Santos IR, Tang J, Schurech M, Kirwan ML, Kopp RE, Zhu K, Li X, Yuan J, Liu W, Li Z (2021). Global blue carbon accumulation in tidal wetlands increases with climate change. National Science Review, 8, nwaa296.

Liu L, Zhu K, Krause SMB, Li S, Wang X, Zhang Z, Shen M, Yang Q, Lian J, Wang X, Ye W, Zhang J (2021). Changes in assembly processes of soil microbial communities during secondary succession in two subtropical forests. Soil Biology and Biochemistry, 154, 108144.

Jones J, Groffman PM, Blair J, Davis FW, Dugan H, Euskirchen ES, Frey SD, Harms T, Hinckley E, Kosmala M, Loberg S, Malone S, Novick K, Record S, Rocha AV, Ruddell B, Stanley EH, Sturtevant C, Thorpe A, White T, Wieder WR, Zhai L, Zhu K (2021). Synergies among environmental science research and monitoring networks: A research agenda. Earth’s Future, 9, e2020EF001631.

Freeman BG, Song Y, Feeley KJ, Zhu K (2021). Montane species track rising temperatures better in the tropics than in the temperate zone. Ecology Letters, 24, 1697-1708.

Clark JS, Andrus R, Aubry-Kientz M, Bergeron Y, Bogdziewicz M, Bragg DC, Brockway D, Cleavitt NL, Cohen S, Courbaud B, Daley R, Das AJ, Dietze M, Fahey TJ, Fer I, Franklin JF, Gehring CA, Gilbert GS, Greenberg CH, Guo Q, Hille Ris Lambers J, Ibanez I, Johnstone J, Kilner CL, Knops J, Koenig WD, Kunstler G, LaMontagne JM, Legg KL, Luongo J, Lutz JA, Macias D, McIntire EJ, Messaoud Y, Moore CM, Moran E, Myers JA, Myers OB, Nunez C, Parmenter R, Pearson S, Poulton-Kamakura R, Ready E, Redmond MD, Reid CD, Rodman KC, Scher CL, Schlesinger WH, Schwantes AM, Shanahan E, Sharma S, Steele M, Stephenson NL, Sutton S, Swenson JJ, Swift M, Veblen TT, Whipple AV, Whitham TG, Wion AP, Zhu K, Zlotin R (2021). Continent-wide tree fecundity driven by indirect climate effects. Nature Communications, 12, 1242.


Zhu K (2020). Understanding forest dynamics by integrating age and environmental change. New Phytologist, 228, 1728-1733.

Wang C, Guan K, Peng B, Chen M, Jiang C, Zeng Y, Wu G, Wang S, Wu J, Yang X, Frankenberg C, Kohler P, Berry J, Bernacchi C, Zhu K, Alden C, Miao G (2020). Satellite footprint data from OCO-2 and TROPOMI reveal significant spatio-temporal and inter-vegetation type variabilities of solar-induced fluorescence yield in the U.S. Midwest. Remote Sensing of Environment, 241, 111728.

Steidinger BS, Bhatnagar JM, Vilgalys R, Taylor JW, Qin C, Zhu K, Bruns TD, Peay KG (2020). Ectomycorrhizal fungal diversity predicted to substantially decline due to climate changes in North American Pinaceae forests. Journal of Biogeography, 47, 772-782.

Qin C, Zhu K, Chiariello NR, Field CB, Peay KG (2020). Fire history and plant community composition outweigh decadal multi-factor global change as drivers of microbial composition in an annual grassland. Journal of Ecology, 108, 611-625.

Moore JR, Argles APK, Zhu K, Huntingford C, Cox PM (2020). Validation of demographic equilibrium theory against tree-size distributions and biomass density in Amazonia. Biogeosciences, 17, 1013–1032.

Liu LZhu K, Wurzburger N, Zhang J (2020). Relationships between plant diversity and soil microbial diversity vary across taxonomic groups and spatial scales. Ecosphere, 11(1):e02999.

Laubmeier AN, Cazelles B, Cuddington K, Erickson KD, Fortin MJ, Ogle K, Wikle CK, Zhu K, Zipkin E (2020). Ecological dynamics: integrating empirical, statistical, and analytical methods. Trends in Ecology and Evolution, 35, 1090–1099.

Lamperty T, Zhu K, Poulsen JR, Dunham AE (2020). Defaunation of large mammals alters understory vegetation and functional importance of invertebrates in an Afrotropical forest. Biological Conservation, 241, 108329.

Jiang F, Zhu K, Cadotte MW, Jin G (2020). Tree mycorrhizal type mediates the strength of negative density dependence in temperate forests. Journal of Ecology, 108, 2601–2610.

He Y, Cheng W, Zhou L, Shao J, Liu H, Zhou H, Zhu K, Zhou X (2020). Soil DOC release and aggregate disruption mediate rhizosphere priming effect on soil C decomposition. Soil Biology and Biochemistry, 144, 107787.

Hakkenberg C, Peet RK, Wentworth TR, Zhu K, Schafale MP (2020). Tree canopy cover constrains the fertility–diversity relationship in plant communities of the southeastern United States. Ecology, 101(10):e03119.

Egerer MH, Wagner B, Lin BB, Kendal D, Zhu K (2020). New methods of spatial analysis in urban gardens inform future vegetation surveying. Landscape Ecology, 35, 761–778.

Compagnoni A, Bibian AJ, Ochocki BM, Levin S, Zhu K, Miller TEX (2020). popler: An R package for extraction and synthesis of population time series from the long-term ecological research (LTER) network. Methods in Ecology and Evolution, 11, 258–264.


Zhu K, Song Y, Qin C (2019). Forest age improves understanding of the global carbon sink. Proceedings of the National Academy of Sciences, 116, 3962-3964.

Yu K, Smith WK, Trugman AT, Condit R, Hubbell SP, Sardans J, Peng C, Zhu K, Peñuelas J, Cailleret M, Levanic T, Gessler A, Schaub M, Ferretti M, Anderegg WRL (2019). Pervasive decreases in living vegetation carbon turnover time across forest climate zones. Proceedings of the National Academy of Sciences, 116, 24662-24667.

Wang C, Zhu K (2019). Misestimation of growing season length due to inaccurate construction of satellite vegetation index time series. IEEE Geoscience and Remote Sensing Letters, 16(8), 1185-1189.

Terborgh J, Zhu K, Álvarez-Loayza P, Cornejo-Valverde F (2019). Seed limitation in an Amazonian floodplain forest. Ecology, 100(5):e02642.

Quan Q, Tian D, Luo Y, Zhang F, Crowther TW, Zhu K, Chen HYH, Zhou Q, Niu S (2019). Water scaling of ecosystem carbon cycle feedback to climate warming. Science Advances, 5:eaav1131.

Mao ZK, Corrales A, Zhu K, Yuan ZQ, Lin F, Ye J, Hao ZQ, Wang XG (2019). Tree mycorrhizal associations mediate soil fertility effects on forest community structure in a temperate forest. New Phytologist, 233, 475–486.

He Y, Zhou X, Cheng W, Zhou L, Zhang G, Zhou G, Liu R, Shao J, Zhu K, Cheng W (2019). Linking improvement of soil structure to soil carbon storage following invasion by a C4 plant Spartina alterniflora. Ecosystems, 22, 859–872.

Chu C, Lutz JA, Král K, Vrška T, Yin X, Myers JA, Abiem I, Alonso A, Bourg N, Burslem DFRP, Cao M, Chapman H, Condit R, Fang S, Fischer GA, Gao L, Hao Z, Hau BCH, He Q, Hector A, Hubbell SP, Jiang M, Jin G, Kenfack D, Lai J, Li B, Li X, Li Y, Lian J, Lin L, Liu Y, Liu Y, Luo Y, Ma K, McShea W, Memiaghe H, Mi X, Ni M, O’Brien MJ, Oliveira AA de, Orwig DA, Parker GG, Qiao X, Ren H, Reynolds G, Sang W, Shen G, Su Z, Sui X, Sun I-F, Tian S, Wang B, Wang X, Wang X, Wang Y, Weiblen GD, Wen S, Xi N, Xiang W, Xu H, Xu K, Ye W, Zhang B, Zhang J, Zhang X, Zhang Y, Zhu K, Zimmerman J, Storch D, Baltzer JL, Anderson‐Teixeira KJ, Mittelbach GG, He F (2019). Direct and indirect effects of climate on richness drive the latitudinal diversity gradient in forest trees. Ecology Letters, 22, 245-255.


Zhu K, Zhang J, Niu S, Chu C, Luo Y (2018). Limits to growth of forest biomass carbon sink under climate change. Nature Communications, 9(1), 2709.

Zhu K, McCormack ML, Lankau RA, Egan JF, Wurzburger N (2018). Association of ectomycorrhizal trees with high carbon-to-nitrogen ratio soils across temperate forests is driven by smaller nitrogen not larger carbon stocks. Journal of Ecology, 106, 524-535.

Wang C, Chen J, Tang Y, Black A, Zhu K (2018). A novel method for removing snow melting-induced fluctuation in GIMMS NDVI3g data for vegetation phenology monitoring: a case study in deciduous forests of North America. IEEE Journal of Selected Topics in Applied Earth Observations and Remote Sensing, 11, 800-807.

Moore J, Zhu K, Huntingford C, Cox P (2018). Equilibrium forest demography explains the distribution of tree sizes across North America. Environmental Research Letters, 13(8), 084019.

Hakkenberg CR, Zhu K, Peet RK, Song C (2018). Mapping multi-scale vascular plant richness in a forest landscape with integrated LiDAR and hyperspectral remote-sensing. Ecology, 99, 474-487.


Wilcox K, Shi Z, Gherardi LA, Lemoine NP, Koerner SE, Hoover DL, Bork E, Byrne K, Cahill J, Collins S, Evans S, Gilgen AK, Holub P, Jiang L, Knapp A, Laura Y, LeCain DR, Liang J, Peñuelas J, Pockman W, Smith M, White S, Zhu K, Luo Y (2017). Asymmetric responses of primary productivity to climate extremes: a synthesis of grassland precipitation manipulations. Global Change Biology, 23, 4376-4385.

Wang C, Chen J, Wu J, Tang Y, Shi P, Black A, Zhu K (2017). A snow-free vegetation index for improved monitoring of vegetation spring green-up date in deciduous ecosystems. Remote Sensing of Environment, 196, 1-12.

Kuang X, Zhu K, Yuan Z, Lin F, Ye J, Wang X, Hao Z (2017). Conspecific density dependence and community structure: insights from 11 years of monitoring in an old-growth temperate forest in northeast China. Ecology and Evolution, 7, 5191-5200.


Zhu K, Chiariello NR, Tobeck T, Fukami T, Field CB (2016). Nonlinear, interacting responses to climate limit grassland production under global change. Proceedings of the National Academy of Sciences, 113, 10589-10594.

Schliep EM, Gelfand AE, Clark JS, Zhu K (2016). Modeling change in forest biomass across the eastern US. Ecological and Environmental Statistics, 23, 23-41.

Ghosh S, Zhu K, Gelfand AE, Clark JS (2016). Joint modeling of climate niches for adult and juvenile trees. Journal of Agricultural, Biological, and Environmental Statistics, 21, 111-130.

Andresen LC, Müller C, Dato G, Dukes J, Emmett BA, Estiarte M, Jentsch-Beierkuhnlein A, Kröel-Dulay G, Luescher A, Niu S, Peñuelas J, Reich P, Reinsch S, Schmidt IK, Schneider M, Sternberg M, Tietema A, Zhu K, Bilton MC (2016). Shifting impacts of climate change: long-term patterns of plant response to elevated CO2, drought, and warming across ecosystems. In: Advances in Ecological Research (eds. Dumbrell, AJ, Kordas, RL, Woodward, G). Academic Press. Oxford, UK. pp. 437-473.

Albright R, Hosfelt J, Kwiatkowski L, Maclaren JK, Mason BM, Nebuchina Y, Ninokawa A, Pongratz J, Ricke KL, Rivlin T, Schneider K, Sesboüé M, Shamberger K, Silverman J, Wolfe K, Zhu K, Caldeira K (2016). Reversal of ocean acidification enhances net coral reef calcification. Nature, 531, 362-365.


Zhu K, Woodall CW, Monteiro JVD, Clark JS (2015). Prevalence and strength of density-dependent tree recruitment. Ecology, 96, 2319-2327.

Woodall CW, Russell MB, Walters BF, D’Amato AW, Zhu K, Saatchi SS (2015). Forest production dynamics along a wood density spectrum in eastern US forests. Trees – Structure and Function, 29, 299-310.

Lankau RA, Zhu K, Ordonez A (2015). Mycorrhizal strategies of tree species correlate with trailing range edge responses to current and past climate change. Ecology, 96, 1451-1458.


Zhu K, Woodall CW, Ghosh S, Gelfand AE, Clark JS (2014). Dual impacts of climate change: forest migration and turnover through life history. Global Change Biology, 20, 251-264.

Terborgh J, Zhu K, Álvarez-Loayza P, Cornejo-Valverde F (2014). How many seeds does it take to make a sapling? Ecology, 95, 991-999.

Clark JS, Gelfand AE, Woodall CW, Zhu K (2014). More than the sum of the parts: forest climate response from joint species distribution models. Ecological Applications, 24, 990-999.

Clark JS, Bell DM, Kwit MC, Zhu K (2014). Competition-interaction landscapes for the joint response of forests to climate change. Global Change Biology, 20, 1979-1991.


Woodall CW, Zhu K, Westfall JA, Oswalt CM, D’Amato AW, Walters BF, Lintz HE (2013). Assessing the stability of tree ranges and influence of disturbance in eastern US forests. Forest Ecology and Management, 291, 172-180.

Woodall CW, Westfall JA, Zhu K, Johnson DJ (2013). Assessing the effect of snow/water obstructions on the measurement of tree seedlings in a large-scale temperate forest inventory. Forestry, 86, 421-427.

Clark JS, Bell DM, Kwit MC, Powell A, Zhu K (2013). Dynamic inverse prediction and sensitivity analysis with high-dimensional responses: application to climate-change vulnerability of biodiversity. Journal of Agricultural, Biological, and Environmental Statistics, 18, 376-404.


Zhu K, Woodall CW, Clark JS (2012). Failure to migrate: lack of tree range expansion in response to climate change. Global Change Biology, 18, 1042-1052.

Zhu K, Ghosh S, Gelfand AE, Clark JS (2012). New approaches to FIA data for understanding distribution, abundance, and response to climate change. In: Moving from Status to Trends: Forest Inventory and Analysis (FIA) Symposium 2012, USDA Forest Service, Northern Research Station, General Technical Report NRS-P-105.

Ghosh S, Gelfand AE, Zhu K, Clark JS (2012). The k-ZIG: flexible modeling for zero-inflated counts. Biometrics, 68, 878-885.

Clark JS, Bell DM, Kwit M, Stine A, Vierra B, Zhu K (2012). Individual-scale inference to anticipate climate-change vulnerability of biodiversity. Philosophical Transactions of the Royal Society B: Biological Sciences, 367, 236-246.


Wang YS, Chu CJ, Zhu K, Shen ZH (2011). Effects of inter-specific variability on biomass allocation: a hierarchical Bayesian approach. Ecological Informatics, 6, 341–344.

Clark JS, Bell DM, Hersh MH, Kwit MC, Moran E, Salk C, Stine A, Valle D, Zhu K (2011). Individual-scale variation, species-scale differences: inference needed to understand diversity. Ecology Letters, 14, 1273-1287.